I have pictures of a frame of standard cell foundation that the bees built Drone cells over the top of.. I tried three times to scrape that foundation and have them re draw it correctly... didn't work, they made drone foundation right back where it was.. I put a new piece of foundation in, and they made more drone foundation, just a bit different in location compared to the old frame..
As I understand it, in a natural hive, the bees want a certain amount of drones, once they reach a specific maturity of hive growth, and they will make drone comb where ever they are capable.. Usually between frames etc.. Natural comb (foundation-less) allows them to satisfy the need.
I have read where beeks get upset and keep ripping it out, and that many think that drone comb is ALL they will draw after a certain point... This has not been the case for me.. Once they satisfy their need, or desire, they return to making normal worker comb.
I do NOT put drone comb in new hives to help them get started, but I will put drone comb from old hives or deadouts in mature hives so that they dont need to BUILD it. I mark those frames for easy recognition when I do inspections.
Understand, this is ONLY what i have observed in my own hives with foundation-less frames.
As I see it, if the hive is big enough to WANT drone comb, then I let them build it and get it over with. When I feed for winter, they normally backfill that drone comb with winter reserves.
Drone comb has advantages as mentioned above. It also puts the drones from YOUR queens out there to mate with any other new queens, hopefully improving the genetics.. of feral hives, neighbors hives that just buy generic packages etc...
Found the pic...